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Fourth Edition

Douglas J. Futuyma and Mark Kirkpatrick

Publication Date - April 2017

ISBN: 9781605356051


In Stock

Retail Price to Students: $189.99

Addresses major themes--including the history of evolution, evolutionary processes, adaptation, and evolution as an explanatory framework--at levels of biological organization ranging from genomes to ecological communities.


Extensively rewritten and reorganized, this new edition of Evolution--featuring a new coauthor: Mark Kirkpatrick (The University of Texas at Austin)--offers additional expertise in evolutionary genetics and genomics, the fastest-developing area of evolutionary biology. Directed toward an undergraduate audience, the text emphasizes the interplay between theory and empirical tests of hypotheses, thus acquainting students with the process of science.

New to this Edition

  • Two authors with broadly overlapping but complementary areas of expertise.
  • Genomic perspectives on evolution are strengthened throughout.
  • The content has a stronger focus on human evolution: an entirely new chapter on the topic (Chapter 21, The Evolutionary Story of Homo sapiens), and new examples throughout the book.
  • Many chapters have been rewritten from the ground up.
  • The book has been entirely reillustrated in a clean, contemporary style that enhances the content.
  • A new Appendix, A Statistics Primer, introduces the concept of a probability distribution, reviews how statistics are used to describe populations, looks at how we estimate quantities, and discusses how hypotheses are tested. It ends with a brief overview of two major frameworks of statistical analysis: likelihood and Bayesian inference. Math is kept to a minimum.


  • The art and illustration program thoroughly enhances the text, updating the visual appeal of the content.
  • Current, comprehensive coverage of evolutionary genetics and genomics.
  • Ideally suited in length and content for one-semester undergraduate evolution courses.

About the Author(s)

Douglas J. Futuyma, Stony Brook University
Mark Kirkpatrick, University of Texas at Austin

Previous Publication Date(s)

March 2013
April 2009
January 2005

Table of Contents

    *Chapters marked by an asterisk have been entirely rewritten.


    Chapter 1. Evolutionary Biology
    Chapter 2. The Tree of Life
    Chapter 3. Natural Selection and Adaptation

    Chapter 4. Mutation and Variation*
    Chapter 5. The Genetical Theory of Natural Selection*
    Chapter 6. Phenotypic Evolution*
    Chapter 7. Genetic Drift: Evolution at Random*
    NEW Chapter 8. Evolution in Space
    Chapter 9. Species and Speciation*

    Chapter 10. All About Sex*
    Chapter 11. How to Be Fit
    Chapter 12. Cooperation and Conflict*
    Chapter 13. Interactions among Species
    Chapter 14. The Evolution of Genes and Genomes*
    Chapter 15. Evolution and Development

    Chapter 16. Phylogeny: The Unity and Diversity of Life
    Chapter 17. The History of Life
    Chapter 18. The Geography of Evolution
    Chapter 19. The Evolution of Biological Diversity
    Chapter 20. Macroevolution: Evolution above the Species Level

    UNIT V. EVOLUTION AND Homo sapiens
    NEW Chapter 21. The Evolutionary Story of Homo sapiens
    Chapter 22. Evolution and Society

    NEW Appendix: A Statistics Primer
    Literature Cited
    Illustration and Photo Credits

Teaching Resources

For Students

Companion Website
The Evolution, Third Edition, Companion Website features review and study tools to help students master the material presented in the textbook. Access to the site is free of charge, and requires no access code. (Instructor registration is required in order for students to access the quizzes.) The site includes the following resources:

* Chapter Outlines and Summaries: Concise overviews of the important topics covered in each chapter.

* Data Analysis Exercises: Expanded for the third edition, these inquiry-based exercises involve students in working with data and analyzing methods and conclusions from published papers.

* Simulation Exercises: Interactive modules that allow students to explore many of the dynamic processes of evolution, and answer questions based on the results they observe.

* Online Quizzes: Quizzes that cover all the major concepts introduced in each chapter. These quizzes are assignable by the instructor.

* Flashcards & Key Terms: Easy-to-use activities that help students learn all the key terminology introduced in each chapter.

* The complete Glossary

For Instructors

Instructor's Resource Library
The Evolution, Third Edition, Instructor's Resource Library includes a variety of resources to help you develop your course and deliver your lectures. The IRL includes the following resources:

* Textbook Figures and Tables: All the figures (including photographs) and tables from the textbook are provided as JPEGs (both high- and low-resolution), reformatted and relabeled for optimal readability when projected.

* PowerPoint Presentations: For each chapter, all of the chapter's figures and tables are provided in a ready-to-use PowerPoint presentation, making it easy to quickly insert figures into your own lecture presentations.

* Answers to the textbook end-of-chapter Problems and Discussion Topics

* Quiz Questions from the Companion Website

* Data Analysis and Simulation Exercises from the Companion Website, with answers

Online Quizzing
A set of online quizzes is available via the Companion Website. These quizzes can be assigned or released for student self-study, at the instructor's discretion. Instructors can also add their own questions to the quizzing system, to create custom quizzes. Results can be viewed online or downloaded for use in gradebook programs. (Instructor registration is required for student access to the quizzes.)

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