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Fifth Edition

Douglas Futuyma and Mark Kirkpatrick

Publication Date - 01 December 2022

ISBN: 9780197619612

702 pages
9 x 10.875 inches

In Stock


The gold standard in undergraduate-level evolutionary biology textbooks.

This new fifth edition presents the field of evolution as a living, breathing science. Extensively revised for clarity and currency, Evolution, 5th Edition, includes updated coverage in evolutionary genetics and genomics to illustrate the rapidly moving science of evolution and emphasizes the interplay between theory and empirical test hypotheses, acquainting students with the process of science. Evolution is available in a dynamic interactive enhanced e-book that allows students hone their problem-solving and data analysis skills while seeing the evolution in the context of their life.

New to this Edition

  • Learning objectives for each chapter.
  • Revised figures to add visual clarity to the key points and include interactivity.
  • The end-of-chapter Problems and Discussion Topics have been completely rewritten. Data analysis questions in appropriate chapters to give students practice in experiential design, and interpreting and analyzing data.
  • Increased coverage of human biology to engage students studying health-related fields.
  • In Chapter 20, extensive coverage of research by 2022 Nobel Prize winner, Svante Pääbo, on the DNA of Neanderthals and Denisovans.
  • "What We Don't Know" boxes convey scientific ideas are constantly being tested and evolving and to help students think like scientists.
  • Discussion of why evolution must embrace the diversity of life rather than focusing on a small number of model systems.
  • Unit VII is a statistics primer that succinctly introduces the concept of a probability distribution, reviews how statistics are used to describe populations, describes how complex datasets are simplified, and discusses how hypotheses are tested.
  • Every chapter is revised and updated.
  • Video and Audio Vignettes.
  • More pedagogical content at the end of each chapter.


  • Ideally suited in length and content for one-semester undergraduate evolution courses.
  • A strong art program enhances the text and visual appeal of the content.
  • Provides an abundance of empirical examples for each major concept.
  • Interactive elements such as self-assessment questions, interactive figures, data exercises with real data sets, simulation exercises, and videos help engage students with the content.
  • Addresses major themes - including the history of evolution, evolutionary processes, adaptation, and evolution as an explanatory framework - at levels of biological organization ranging from genomes to ecological communities
  • Chapters build on each other in an easy-to-understand progression.

About the Author(s)

Douglas Futuyma is recognized as one of the world's leading evolutionary biologists. His research is focused on the evolution and coevolution of ecological interactions among different species, especially between plants and herbivorous insects. His work with insects and host plants began as research to account for all of the genetic variation that has occurred in evolution. He is well known as the author of a leading college textbook on biological evolution and of a trade book that addresses creationist opposition to teaching avolution.

Mark Kirkpatrick is T.S. Painter Centennial Professor of Genetics at the University of Texas, Austin and heads the Kirkpatrick Lab. His work uses mathematical models and develops statistical tools to test hypotheses. Topics of focus include: chromosome evolution, speciation, sexual selection and species ranges.

Table of Contents

    Chapter 1: An Idea that Changed the World
    Chapter 2: The Tree of Life
    Chapter 3: Natural Selection and Adaptation
    Chapter 4: Mutation and Variation
    Chapter 5: The Genetical Theory of Natural Selection
    Chapter 6: Genetic Drift: Evolution at Random
    Chapter 7: Quantitative Traits
    Chapter 8: Evolution in Space
    Chapter 9: Species and Speciation
    Chapter 10: The Evolution of Genes and Genomes
    Chapter 11: Evolution and Development
    Chapter 12: All about Sex
    Chapter 13: Cooperation and Conflict
    Chapter 14: Evolutionary Ecology: How to be Fit
    Chapter 15: Interactions among Species
    Chapter 16: Phylogeny: The Unity and Diversity of Life
    Chapter 17: The History of Life
    Chapter 18: The Geography of Evolution
    Chapter 19: Macroevolution: Evolution above the Species Level
    Chapter 20: The Evolutionary Story of Homo sapiens
    Chapter 21: Evolution and Society
    Chapter 22: A Statistics Primer

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